When Kratos was on his mission to stop Ares' reign of terror, he was told that he would need the power held within Pandora's Box to defeat him. He used the skulls for their purpose, leaving the bodies without skulls. The Temple was then left without anything to stop mortals from reaching the box (including Kratos).
- In the Cliffs of Madness, several deceased soldiers' bodies are found. It remains unknown how those soldiers got so far without having taken the skulls from the sons' corpses. It is possible that this was a mistake overlooked by the developers, although it's possible that they have conquered the lower floors but died as a result in the upper floors as the Cliffs of Madness is very unforgiving as Pathos called it a "Maze with no true way out"